Mushroom Trivia
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gill spacing

Terms discussed: 2:1, 4:1, close, crowded, distant, extremely crowded, extremely distant, subdistant




Photo of Amanita cokeri by Leon Shernoff
Amanita cokeri
Gill spacing can be a useful character in determining a mushroom's identity, but it does have the drawback that we don't have a term for "normal", or "average".
The gill spacing shown here, for instance, is probably the most usual that you will see (also in store-bought mushrooms, for example); it is considered crowded, or close.

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Image of Lactarius glaucescens from Eugen Gramberg (1913) Pilze unserer Heimat
Lactarius glaucescens
Extremely crowded gills are seen in the Peppery section of Lactarius, where the gills are so close together that they sometimes seem like a series of ridges on the surface of a solid mass.

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Image of Hygrophorus fuligineus from Eugen Gramberg (1913) Pilze unserer Heimat
Hygrophorus fuligineus
Subdistant gills are not quite distant, but not close either. Some people use the word close for this sort of gill spacing.
Once your gills get subdistant or farther apart, it's common to find partial gills, or lamellulae. Partial gills are those that only reach part of the way from the cap to the stem. They are usually present in one of two ratios. In the Hygrophorus picture, we have a 4:1 ration: between each full gill, there is a partial gill that reaches about half way to the stalk, with two tiny partial gills on either side of it. Some fungi have a 2:1 ratio, where there is only one partial gill between each full gill.

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Image of Camarophyllus pratensis from Eugen Gramberg (1913) Pilze unserer Heimat
Camarophyllus pratensis
These gills are distant. Extremely distant gills can actually be counted easily: there's usually just a dozen to a dozen and a half for the whole cap.

Of course, none of this has really been standardized. " Close", in particular, has been used in different publications to mean what I call crowded, what I call subdistant, or to mean something in between. While it has an "obvious" natural meaning, I try not to use it (it may slip out a few times during the course of the web site), as I find it just ambiguous, unless specially defined: for example, Lincoff (1987) gives explicit pictures of only three states: crowded, close, and distant. He may very well have the right idea.

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